The species of the Melitaea ala Staudinger, 1881 complex are distributed in Central Asia. Here we show that this complex is a monophyletic group including the species, M. ala, M. kotshubeji Sheljuzhko, 1929 and M. enarea Fruhstorfer, 1917. The haploid chromosome number n=29 is found in M. ala and M. kotshubeji and is, most likely, a symplesiomorphy of the M. ala complex. We show that M. ala consists of four subspecies: M. ala zaisana Lukhtanov, 1999 (=M. ala irtyshica Lukhtanov, 1999, syn. nov.) (South Altai, Zaisan Lake valley), M. ala ala (Dzhungarian Alatau), M. ala bicolor Seitz, 1908 (North, East, Central and West Tian-Shan) and M. ala determinata Bryk, 1940 (described from “Fu-Shu-Shi”, China). We demonstrate that M. kotshubeji kotshubeji (Peter the Great Mts in Tajikistan) and M. kotshubeji bundeli Kolesnichenko, 1999 (Alai Mts in Tajikistan and Kyrgyzstan) are distinct taxa despite their geographic proximity in East Tajikistan. Melitaea enarea is widely distributed in the southern part of Central Asia and is sympatric with M. kotshubeji.

中文翻译：

---

Melitaea ala Staudinger, 1881 物种复合体（鳞翅目，蛱蝶科）的染色体和 DNA 条形码分析

Melitaea ala Staudinger, 1881 复合体的物种分布于中亚。在这里，我们表明该复合体是一个单系群，包括物种 M. ala、M. kotshubeji Sheljuzhko，1929 和 M. enarea Fruhstorfer，1917。在 M. ala 和 M. kotshubeji 中发现单倍体染色体数 n=29很可能是 M. ala 复合体的共形同形。我们表明 M. ala 由四个亚种组成： M. ala zaisana Lukhtanov, 1999 (= M. ala irtyshica Lukhtanov, 1999, syn. nov.) (South Altai, Zaisan Lake Valley), M. ala ala (Dzhungarian Alatau) , M. ala bicolor Seitz, 1908 (North, East, Central and West Tian-Shan) and M. ala determinata Bryk, 1940 (描述自“Fu-Shu-Shi”, China)。我们证明了 M. kotshubeji kotshubeji（塔吉克斯坦的彼得大帝山）和 M. kotshubeji bundeli Kolesnichenko，1999（塔吉克斯坦和吉尔吉斯斯坦的 Alai Mts）是独特的分类群，尽管它们在东塔吉克斯坦的地理上很接近。Melitaea enarea广泛分布于中亚南部，与M. kotshubeji同域分布。