Bryozoan–serpulid–algal–thrombolite bioherms of up to 50 cm size are described from the Sarmatian (upper Middle Miocene) of the Central Paratethys. They occur on top of lower Sarmatian carbonate sediments of high-energy conditions and the individual bioherms settle on crests of ripples. The buildups are overlain and partly truncated by cross-bedded oolites of late Sarmatian age. Buildup growth starts with a Cryptosula/Hydroides (bryozoan/serpulid) pioneer community, followed by nodular Schizoporella (bryozoan) colonies overgrown by coralline algae/microbial mats and a thrombolite with calcareous algal filaments. All these constituents form a framestone fabric which is overall dominated by bryozoans labeling them as bryoherms. Inside the bioherms ecological successions of higher frequencies occur which are interpreted to reflect short-time environmental fluctuations such as nutrient availability, oxygenation (possible anoxia), salinity (possible brackish water), temperature and water level. The internal succession in individual bioherms is related to long-term environmental changes including general shallowing, increasing nutrient supply and decreasing water circulation and oxygenation. The described bioherms are most similar to modern bryostromatolites of the Coorong lagoon in S Australia and also similar to structures in the Netherlands. The widespread occurrence of bryoherms/bryostromatolites in the Central Paratethys suggests a phase of considerable eutrophication during the early Sarmatian.

最大 50 厘米大小的苔藓虫-蛇纹石-藻类-凝块岩生物礁描述于中副提斯的萨尔马提亚阶（上中新世）。它们出现在高能条件下萨尔马提亚碳酸盐沉积物的顶部，单个生物礁落在涟漪的波峰上。这些岩堆被萨尔马提亚晚期的交错层状鲕岩覆盖并部分被截断。累积生长始于隐孢子虫/水螅虫（苔藓动物/蛇类）先驱群落，然后是结节裂隙菌（苔藓虫）菌落长满珊瑚藻/微生物垫和含钙质藻丝的凝块石。所有这些成分形成了一个框架结构，该结构总体上由苔藓虫主导，将它们标记为苔藓虫。在生物礁内部，发生了更高频率的生态演替，这被解释为反映了短时间的环境波动，例如养分可用性、氧合作用（可能缺氧）、盐度（可能是微咸水）、温度和水位。个别生物礁的内部演替与长期环境变化有关，包括普遍变浅、营养供应增加以及水循环和氧合作用减少。所描述的生物礁与澳大利亚南部 Coorong 泻湖的现代苔藓叠层石最相似，也与荷兰的结构相似。