The mixed siliciclastic–carbonate Teniet El Klakh Formation (Middle Jurassic, Bajocian) mainly consists of a monotonous, clay and fine-grained sandstone alternation with some intercalated carbonate beds. The trace fossil record displays moderate to high ichnological diversity with different behaviours, consisting of 28 ichnotaxa belonging to 23 ichnogenera: isp., , , isp., , , , isp., , , isp., cf. , , cf. , cf. , isp., isp., cf. , , isp., isp., , isp., var. , , , cf. , and . The distribution of these trace fossils in space and time is controlled by various environmental features, mostly referred to the accumulation of organic matter in softground substrate during short phases of low hydrodynamics in well-oxygenated environments, thus allowing the occurrence of abundant and diverse invertebrates frequently dominated by deposit-feeders. The integrated sedimentological and ichnological features suggest that the Teniet El Klakh Formation had been deposited in a storm-influenced continental shelf setting ranging from the lower offshore to the lower shoreface zones with fluctuating energy regime, corresponding to the archetypal and proximal ichnofacies transitional to the ichnofacies. The ichnological assemblage described herein is original and rather unique, since most of trace fossils are reported herein for the first time from the Ksour Mountains.

中文翻译：

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来自中侏罗世（巴约世）Teniet El Klakh 组（阿尔及利亚西撒哈拉阿特拉斯）的多样化遗迹化石组合

混合硅质碎屑-碳酸盐岩 Teniet El Klakh 组（中侏罗世，巴约世）主要由单调的粘土和细粒砂岩互层组成，并夹有一些碳酸盐岩层。痕迹化石记录显示出中等至高度的具有不同行为的物种多样性，由属于 23 个物种属的 28 个物种组成：isp., , , isp., , , , isp., , , isp., cf. , , 参见，参见, isp., isp., cf. , , isp., isp., , isp., var. , , , 参见， 和 。这些痕迹化石在空间和时间上的分布受到各种环境特征的控制，大多是指在充氧良好的环境中，在短暂的低水动力阶段，有机质在软土基质中的积累，从而使得丰富多样的无脊椎动物频繁出现。以存款供给者为主。综合沉积学和地貌学特征表明，Teniet El Klakh 组沉积在风暴影响的大陆架环境中，范围从近海下部到下岸面区域，能量状态波动，对应于过渡到地层相的原型和近端地层相。 。本文描述的技术组合是原始且相当独特的，因为本文中的大多数痕迹化石是首次从库苏尔山脉报道。