Conodonts are a group of extinct marine organisms widely used in Paleozoic and Triassic biostratigraphy. The genus Siphonodella comprises ozarkodinid conodonts from the latest Famennian to the middle Tournaisian. The phylogenetic model of this genus plays an important role in biostratigraphy. This study is based on the authors' and published data on 31 species of siphonodellids of cosmopolitan and East European groups. A new phylomorphogeny of Siphonodella is proposed on the basis of known mechanisms of morphological transformations of siphonodellid P₁ elements during speciation processes, presence of interspecies transitional forms, and data on ontogeny. The main morphogenetic pathways in the evolution of siphonodellids are the following: formation and transformation of the rostrum, transformation of the platform outline, transformation of the platform ornamentation, transformation of the keel, and histological transformations. In the process of modeling the phylomorphogeny of Siphonodella, some taxonomic problems are solved: a new species S. thompfelli is described, a new variant of the division of S. lobata (Branson and Mehl) into morphotypes is proposed, the validity of S. plana (Huddle), S. praeplana Bardasheva, Bardashev, Weddige and Ziegler, and S. gladia Zhuravlev and Plotitsyn is clarified, and the knowledge of the morphospace of most representatives of the genus is extended. The phylogeny of siphonodellids provides a basis for the new phylogenetic zonal sequences based on the conodont phylogenetic-zone concept.

牙形刺是一组广泛应用于古生代和三叠纪生物地层学的已灭绝海洋生物。管管藻属(Siphonodella)包括从法门纪晚期到图尔奈中期中期的 ozarkodinid 牙形刺。该属的系统发育模型在生物地层学中发挥着重要作用。这项研究基于作者和已发表的关于世界性和东欧群体的 31 种管水动物的数据。基于已知的管水藻形态转变机制，提出了一种新的管水藻系统形态发生学 P ₁物种形成过程中的元素、种间过渡形式的存在以及个体发育数据。管类动物进化的主要形态发生途径有：吻部的形成和改造、平台轮廓的改造、平台装饰的改造、龙骨的改造和组织学改造。在对管管藻系统形态发生进行建模的过程中，解决了一些分类学问题：描述了一个新种S. thompfelli ，提出了管管藻 (S. lobata)形态型划分的新变体（Branson 和 Mehl）， S. thompfelli的有效性。plana (Huddle)、S. praeplana Bardasheva、Bardashev、Weddige 和 Ziegler，以及S.gladia Zhuravlev 和 Plotitsyn 得到澄清，并且扩展了该属大多数代表的形态空间知识。管管科动物的系统发育为基于牙形体系统发育区概念的新系统发育区序列提供了基础。