The hard tick (Ixodida: Ixodidae) from the mid-Cretaceous (ca. 100 Ma) Burmese amber of Myanmar was first described in a new, extinct genus based on a larva. Here, several more larvae are documented providing additional morphological data based on optical studies using a Keyence VHX-6000 Digital Microscope supplemented by computed microtomography (µ-CT). We confirm the presence of 14 festoons (not 13) on the posterior margin of the body, a character unique for the genus. We further describe an unusual pattern of hypostome dentition in the mouthparts, with an unequal and uneven 2/2 arrangement. Moreover, on the internal side of the palps, to the second and third segments five tooth-like elements are present, similar to the hypostome dentition. Fossil ticks are generally very rare, thus the discovery of numerous new specimens of suggests that it may have been relatively common in the Burmese amber forest. Larval instars predominate in the fossil record and we might speculate that they preferred arboricolous hosts.

中文翻译：

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缅甸琥珀中已灭绝的硬蜱 Compluriscutula vetulum（蛛形纲：Ixodida）幼虫的新记录，及其形态注释

来自白垩纪中期（约 100 Ma）缅甸琥珀的硬蜱（Ixodida：Ixodidae）首次在基于幼虫的新灭绝属中被描述。此处，记录了更多幼虫，使用基恩士 VHX-6000 数码显微镜并辅以计算机显微断层扫描 (μ-CT) 进行光学研究，提供了额外的形态学数据。我们确认在身体后缘存在 14 个花彩（不是 13 个），这是该属独有的特征。我们进一步描述了口器中不寻常的下口牙列模式，具有不等且不均匀的 2/2 排列。此外，在触须的内侧，到第二节和第三节存在五个齿状元件，类似于下口齿列。蜱虫化石通常非常罕见，因此大量新标本的发现表明，蜱虫化石在缅甸琥珀森林中可能相对常见。幼虫龄在化石记录中占主导地位，我们可能推测它们更喜欢树栖寄主。