Thomas Cavalier-Smith, born in London, U.K., on October 21, 1942, was a Professor of Evolutionary Biology in the Department of Zoology at the University of Oxford at the time of his death on March 19, 2021. Credited with at least 235 research works and over 20,000 citations, Cavalier-Smith was a well-known and widely respected scientist who took a bold and detailed approach to understanding major transitions in evolution, including the role of endosymbiosis. He was noted for his willingness to question theories and constantly accumulate and evaluate data, motivated by science for the sake of science. This paper reviews Thomas Cavalier-Smith's major accomplishments, examines his theoretical approaches, and provides highlights from the “Tree of Life Symposium” sponsored by the International Society of Protistologists (ISOP) and the International Society of Evolutionary Protistology (ISEP) on June 21, 2021, to celebrate Tom's life and work.

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悼念：托马斯·卡瓦利尔·史密斯 (1942–2021)

托马斯·卡瓦利尔·史密斯 (Thomas Cavalier-Smith)，1942 年 10 月 21 日出生于英国伦敦，于 2021 年 3 月 19 日去世时是牛津大学动物学系进化生物学教授。被认为至少有 235 Cavalier-Smith 的研究著作和超过 20,000 次引用，是一位广受尊敬的著名科学家，他采用大胆而详细的方法来理解进化中的重大转变，包括内共生的作用。他以愿意质疑理论并不断积累和评估数据而闻名，他的动机是为了科学而科学。本文回顾了托马斯·卡瓦利尔·史密斯 (Thomas Cavalier-Smith) 的主要成就，检验了他的理论方法，并提供了国际原生动物学会 (ISOP) 和国际进化原生动物学会 (ISEP) 于 6 月 21 日主办的“生命之树研讨会”的要点， 2021 年，庆祝汤姆的生活和工作。