We studied a collection of the Silurian (Rhuddanian) multi-branched dendroids, including reticulate, benthic dendroid graptolites of the Callograptidae Hopkinson (in Hopkinson and Lapworth, 1875), from the Gorny Altai, Russia. Species of Dictyonema Hall, Koremagraptus Bulman, Callograptus Hall, and Desmograptus Hopkinson were identified and described. The graptolites are represented by in situ burial of colonies with 3D preservation of tubaria (= rhabdosomes). Complete (intact) colonies of dendroid graptolites are characterized by successive development stages: juvenile, neanic, ephebic (adults), while the general configuration of the tubarium changed from the proximal to distal. In a number of cases, later stage of colony development — the gerontic stage (senilis) is found. We carried out morphological analysis of dendroid graptolite tubaria (Callograptidae) of various development stages, showing eight morphotypes of the colonies, namely, the cylinder, disc, disc on a stem, hemisphere, cone, wineglass, blooming flower, and funnel type.

我们研究了一系列志留纪（Rhuddanian）多分支树状体，包括来自俄罗斯戈尔诺阿尔泰地区的 Callograptidae 霍普金森（Hopkinson 和 Lapworth，1875 年）的网状底栖树状笔石。鉴定并描述了 Dictyonema Hall、Koremagraptus Bulman、Callograptus Hall 和 Desmograptus Hopkinson 的物种。笔石以原位埋藏的菌落和 3D 保存的管状体（=弹状体）为代表。完整（完整）的树状笔石群落的特征是连续的发育阶段：幼年期、幼年期、幼年期（成体），而管状体的总体构型从近端到远端发生变化。在许多情况下，发现了群体发育的后期阶段——老年期（senilis）。我们对不同发育阶段的树状笔石管状体（Callograptidae）进行了形态学分析，发现了其菌落的8种形态类型，即圆柱型、盘型、茎盘型、半球型、圆锥型、酒杯型、盛开的花型和漏斗型。