Consistent species identification is foundational to biological research and requires coordination among a diversity of researchers and institutions. However, such consistency may be hindered for rare organisms where specimens, identification resources, and taxonomic experts are few. This is often the case for deep-sea taxonomic groups. For example, the deep-sea gastropod genus Provanna Dall, 1918 is common at chemosynthetic sites throughout the world’s oceans, yet no formal guide to these species has yet been produced. Recent exploration has recovered new specimens of Provanna from hydrocarbon seeps off the Pacific Costa Rica Margin. The current work assesses the species identity of these specimens using shell morphology, radular morphology, and genetic barcoding (mitochondrial CO1 and nuclear H3). Records of occurrence for P. laevis Warén & Ponder, 1991, P. ios Warén & Bouchet, 1986, and P. pacifica Warén & Bouchet, 1986 are herein presented from the Costa Rica Margin. A critical taxonomic review of the 29 extant species within this genus was conducted and their genetic, morphological, and biogeographical distinction assessed. In this review, genetic and morphological support was found for nearly all current species delineations except for P. glabraOkutani et al., 1992, syn. nov. and P. laevis, syn. nov., which are herein synonymized to P. laevis, and for P. ios, syn. nov. and P. goniata Warén & Bouchet, 1986, syn. nov., which are synonymized to P. ios. Finally, the first species identification key for the extant species in this genus is presented. This work clarifies the taxonomy and systematics of this deep-sea gastropod genus and contributes a novel polytomous key for use in future research.

中文翻译：

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哥斯达黎加边缘的 Provanna（腹足纲、Provannidae）新记录以及该属的识别码

 一致的物种鉴定是生物学研究的基础，需要各种研究人员和机构之间的协调。然而，对于标本、鉴定资源和分类学专家很少的稀有生物来说，这种一致性可能会受到阻碍。深海分类群通常就是这种情况。例如，深海腹足动物属 Provanna Dall, 1918 在世界各地海洋的化学合成地点很常见，但尚未制定这些物种的正式指南。最近的勘探从太平洋哥斯达黎加边缘的碳氢化合物渗漏中发现了新的普罗瓦纳样本。目前的工作使用壳形态、齿状形态和遗传条形​​码（线粒体 CO1 和核 H3）评估这些标本的物种身份。本文介绍了哥斯达黎加边缘的 P. laevis Warén & Ponder, 1991、P. ios Warén & Bouchet, 1986 和 P. pacifica Warén & Bouchet, 1986 的出现记录。对该属内 29 个现存物种进行了严格的分类学审查，并评估了它们的遗传、形态和生物地理差异。在这篇综述中，除了 P. glabraOkutani 等人，1992，syn. 之外，几乎所有当前物种的描述都发现了遗传和形态学支持。十一月 和 P. laevis，syn。nov.，其在本文中与P. laevis同义，对于P. ios同义。十一月 和 P. goniata Warén & Bouchet，1986，syn。nov.，它们是 P.ios 的同义词。最后，提出了该属现存物种的第一个物种鉴定关键码。这项工作阐明了这种深海腹足动物属的分类学和系统学，并为未来的研究提供了一种新颖的多节键。