Abstract

A new classification system for Protozygoptera is proposed. It is based on mass material from the Isady, Chekarda and Soyana localities. The new system is based on a set of wing venation characters. The hypothesis on the division of Protozygoptera based on their body structures is supplemented and confirmed (Nel et al., 2012). A three-segmented tarsal structure has been established for the archizygopteran family Kennedyidae. The infraorder Permagrionomorpha infraordo nov. is established. An additional character has been proposed to separate archizygopterans and permagrionomorphs based on the structure and morphology of the thorax. Lodeviidae is synonymized with Permagrionidae (junior synonym). Two new families, four new genera and 14 new species are described. A new family Progoneuridae fam. nov. is described based on a significant reduction in the wing longitudinal and transverse venation. It includes the genera Progoneura Carpenter, 1931; Azaroneura Nel et al., 2012; Progophlebia Felker, 2021, and Issadoneura gen. nov. (with a new species Issadoneura marilevorum gen. et sp. nov.). Based on re-examination of the type material from Chekarda, the genus Engellestes Nel et al., 2012 is assigned to a new family Engellestidae fam. nov. The following voltzialestids are described from the Upper Permian of the Vologda Region: Prototerskeja dubia gen. et sp. nov., Permosticta elegans gen. et. sp. nov., and Permosticta parva gen. et sp. nov. The diagnosis of the family Voltzialestidae is changed because of the inclusion of new genera. The genus Sushkinia Martynov, 1930 is transferred to Permagrionidae. The following permagrionids are described: Permagrion sharovi sp. nov, Permolestes obscurus sp. nov., P. vjatkensis sp. nov., Salagoulestes martynovi sp. nov., Solikamptilon aequus sp. nov., Sushkinia angulata sp. nov., Triadolestes sakmarensis gen. et sp. nov., and T. mutovkensis gen. et sp. nov. The new material from Isady and Kargala suggests closer relationships of the Permian Permagrionomorpha and Triassic Triadophlebiomorpha. In the family Kennedyidae, Kennedya sylvensis sp. nov. and K. volatica sp. nov. are described from Chekarda, and K. ivensis Nel et al., 2012 is redescribed from Soyana.

中文翻译：

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东欧地台二叠纪豆娘新类群（原接合翅目）

摘要

提出了一种新的原接合翅目分类系统。它基于来自伊萨迪、切卡达和索亚纳地区的大量材料。新系统基于一组翼脉特征。原接合翅目根据身体结构进行划分的假说得到补充和证实(Nel et al., 2012)。已为祖翅目肯尼迪科建立了三节跗骨结构。下目 Permagrionomorpha infraordo nov.成立。根据胸部的结构和形态，提出了一个额外的特征来区分Archizygopterans和Permagrionomorphs。 Lodeviidae 与 Permagrionidae （初级同义词）同义。描述了两个新科、四个新属和 14 个新种。原神经科一新科。十一月是基于机翼纵向和横向脉络的显着减少来描述的。它包括Progoneura Carpenter属，1931；阿扎罗内拉内尔等人，2012；Progophlebia Felker，2021，和Issadoneura gen。十一月（有一个新物种Issadoneura marilevorum gen. et sp. nov.）。根据对 Chekarda 模式材料的重新检查，Engellestes Nel 等人于 2012 年将其归属于一个新的 Engelestidae 科。十一月以下 voltzialestids 是在沃洛格达地区上二叠统描述的：Prototerskeja dubia gen.等十一月，Permosticta elegans gen。等sp。 nov. 和Permosticta parva gen.等十一月由于纳入了新属，Voltzialestidae 科的诊断发生了变化。Sushkinia Martynov属，1930 年被转移到 Permagrionidae 科。描述了以下 Permagrionids：Permagrion sharovi sp。十一月，Permolestes obscurus sp。十一月, P. vjatkensis sp。十一月，Salagoulestes martynovi sp。十一月，Solikamptilon aequus sp。十一月，Sushkinia angulata sp。十一月，Triadolestes sakmarensis gen.等nov. 和T. mutovkensis gen.等十一月来自伊萨迪和卡尔加拉的新材料表明二叠纪 Permagrionomorpha 和三叠纪 Triadophlebiomorpha 之间的关系更为密切。在肯尼迪科 (Kennedyidae) 中，肯尼迪 (Kennedya sylvensis sp)。十一月和K. volatica sp。十一月是从 Chekarda 中描述的，K. ivensis Nel 等人，2012 从 Soyana 中重新描述的。