The current study focuses on the emblematic / species complex (tribe Lemmini) in the European Pleistocene fossil record. The members of the two genera occupy distinct ecological niches and have different external appearances, but they are remarkably similar in their dental morphology, so that they were commonly thought of as undistinguishable in the fossil record. Thus, more or less all European Lemmini fossils have been assigned to the genus . In the Early Pleistocene site of Schernfeld (Germany), the species had been described. It was thought by some authors that all Lemmini from Early to late Middle Pleistocene belong to this species.

中文翻译：

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对西欧和中欧早更新世和中更新世的莱米尼化石标本的重新研究：进化和古环境的影响

目前的研究重点是欧洲更新世化石记录中的标志性/物种复合体（莱米尼部落）。这两个属的成员占据着不同的生态位，具有不同的外观，但它们的牙齿形态却非常相似，因此在化石记录中通常认为它们无法区分。因此，或多或少所有欧洲莱米尼化石都被归入该属。在德国舍恩菲尔德的早更新世遗址中，该物种已被描述。一些作者认为，中更新世早到晚的所有 Lemmini 都属于这个物种。